Food and Agriculture Organization of the United Nationsfor a world without hunger. Pakistan Study Report. Cast nets are often used for partial harvesting in small and backyard ponds. Here induced breeding by hormones method is briefly described. Mystus seenghala 7. Complete harvesting is possible only through draining. The spawn recovery of mrigal usually ranges from 100 000 to 150 000/kg. Cirrhinus mrigala is a species of ray-finned fish in the genus Cirrhinus.It is found in northern India, Pakistan, Bangladesh and Nepal. These exotic species are also occasionally employed in polyculture along with the Indian major carp species. Singh, D.M. Pectoral fin… Asian Fishery Society Indian Branch, Mangalore, India. During recent years a small quantity of Indian major carps is being exported from India to the Middle East, degutted and frozen. Ltd., New Delhi, India. The Lao People's Democratic Republics, Thailand, Vietnam, Pakistan, Myanmar and Nepal also use mrigal as one of the principal components in carp polyculture systems. Mass production of intergeneric hybrid catla (Catla catla x Labeo rohita) and its growth in ponds, small and large reservoirs of Madhya Pradesh. © Copyright 2020 CAB International. Shams works in freshwater ecosystems, primarily on fish diversity in terms of their availability, and richness; he is also interested in aquatic invasive species and … 93245). The main objective of this study was to determine the acute pathological effect of barium chloride. FAO Fisheries Synopsis No. Utilization of Cirrhinus Mrigala Fingerlings. Manila, Asian Development Bank & Bangkok, Network of Aquaculture Centres in Asia-Pacific. & Ayyappan, S. 1998a. Khoke, U.C. To reduce feed cost carbohydrates are utilized in fish feed formulation because carbohydrates are most economical, low cost than lipids and protein and a rich source of energy. Central Institute of Freshwater Aquaculture, India. 541 pp. Fish-Pak: a dataset of 6 different fish species i.e. Nursery rearing of Indian major carp fry under different stocking densities. In: Bangladesh Study Report. & Varghese, T.J. 1980. Cirrhina leschenaultii (Valenciennes, 1842), Â Â Â Â Â Â Â Â Â Â Â Â Subphylum: Vertebrata, Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Class: Actinopterygii, Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Order: Cypriniformes, Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Family: Cyprinidae, Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Genus: Cirrhinus, Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Species: Cirrhinus cirrhosus. Rome, FAO. Cirrhinus mrigala did equally well on all ingredients while Hypophthalmichthys molitrix did very poorly. ISOLATED FROM SKIN ULCERS OF CIRRHINUS MRIGALA Parvati Sharma, R.C. classification of freshwater fishes of the madduvalasa reservoir in the palakonda division of srikakulam district and their status was evaluated. Eect of chromium on corn digestibility in Cirrhinus mrigala in the absence of natural feed Cirrhinus mrigala feeds on artificial feed (Khan et al., 2015). In: Regional Study and Workshop on Aquaculture sustainability and the Environment. Clarias Batrachus 9. Fish and Fisheries of India. 90 pp. Channa gachua […] 1968. C. cirrhosus has four barbells whereas mrigala has only two barbells; dorsal branched rays are 15-16 in cirrhosus vs. 12-13 in mrigala.” From Simonsen et al. Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students.ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Nepal Study Report. Manila, Asian Development Bank & Bangkok, Network of Aquaculture Centres in Asia-Pacific. Histological analysis demonstrated that the pancreatic tissues were dispersed within the hepatic parenchyma as a hepatopancreas in C. mrigala and O. nilotica. In English, it is called Mrigal Carp, or Mrigal… The classification results indicate that only the DAGMSVM meets the requirement of time efficiency for the system. Without molecular data, it is very difficult to identify new or existing species. Choudhury, S.N. Manila, Asian Development Bank & Bangkok, Network of Aquaculture Centres in Asia-Pacific. Handbook of Zoological Survey of India, Volume 2. The grow-out culture of mrigal in polyculture systems is confined to earthen ponds and normal management practice includes predatory and weed fish control with chemicals or plant derivatives; stocking of fingerlings at a combined density of 4 000-10 000 fingerlings/ha; fertilization with organic manures like cattle dung or poultry droppings and inorganic fertilizers; supplementary feeding with a mixture of rice bran/wheat bran and oil cake; and fish health monitoring and environmental management. Taxonomic Notes: Cirrhinus reba was originally described as Cyprinus reba by Hamilton (1822) from Bengal and Behar (Bihar), India. Journal of Aquaculture in the Tropics, 13:143-150. Adults are bottom dwellers.It is an illiophage in its feeding habit and stenophagous; detritus and decayed vegetation form its principal food components, while phytoplankton and zooplankton comprise the rest. Shrimp and carp aquaculture and the environment. 1981. CABI is a registered EU trademark. • Large cycloids scales presents • Body dark grey on back and silvery on the belly sides • One pair of short barbles present on the mouth. In: Regional Study and Workshop on Aquaculture sustainability and the Environment. 1995. Four experimental diets … Aquaculture products can anchorage several pathogenic bacterial species, which are known as part of the micro-flora of the environment. 1998b. ), Proceedings of the workshop on carp seed production technology, pp. In: T. J. Pandian (ed. Improvements in feeding and health management. It is also called Migel or Mirka. [Draft report], Pathak, S.C. & Palanisamy, K. 1995. The species is mostly marketed fresh in local markets. Post-harvest processing and value-addition is almost non-existent at present in any of the producing countries. The bottom dwelling habit of mrigal hinders its effective harvesting by dragnet, the most common gear used in carp culture. CIFA. However, being a bottom dweller, the harvesting of this species is a perpetual problem, especially in undrainable ponds. The Mrigal fish can grow very big. As mrigal does not breed in confined waters, injections of pituitary extract and other synthetic commercial formulations of purified salmon gonadotropin and dopamine antagonists such as Ovaprim, Ovatide and Wova-FH have also been successfully used in recent years. Indian Journal of Fisheries, 45(2): 163-168. This species has long been important in polyculture with other native species, mainly in India. However, consumer preference makes the prices obtained for these species always higher than those for common carp and Chinese carps. Cirrhina mrigala Day, 1878. Mrigal is normally cultured along with the other two Indian major carps - catla (, Mass scale seed production of mrigal in hatcheries through induced breeding now supplies almost the entire seed requirement in all the producing countries, although riverine collection still forms the source of seed in certain small areas. The Chinese circular hatchery is the most common system used. [Draft report]. Detailed coverage of invasive species threatening livelihoods and the environment worldwide. Jhingran, V.G. India has vast freshwater resources of approximately 5.4 million hectares in the form of ponds, tanks, lakes and reservoirs. Isolation and Identification of Pathogenic Bacteria Associated with Diseased Fish Cirrhinus mrigala (Ham) in Glass Aquarium. Improvements in induced breeding and seed production, which have removed the reliance on the capture of natural riverine seed. A first report on quadruple spawning of Catla catla (Ham.). The gonadotropin hormone (FSH and LH) secreted by pituitary gland influences the maturation of gonads and spawning in fishes. Mrigal (Cirrhinus mrigala), a carp endemic to Indo-Gangetic riverine systems, is one of the three Indian major carp species cultivated widely in Southeast Asian countries. Being a low input-based system, carp culture has not generally been perceived as a threat to the environment. Regional Study and Workshop on Aquaculture Sustainability and the Environment. May 2011: Ref. 41236).Please send references, or more studies are needed. Annual Report Central Institute of Freshwater Aquaculture. σ. Abstract- An eight week feeding trial was conducted to evaluate the potential of Gliricidia maculata leaf meal as dietary protein source in the diet of Cirrhinus mrigala fingerlings. The grow-out period is usually one year, during which mrigal grows to about 600-700 g. Production is normally 3-5 tonnes/ha/yr, with mrigal contributing about 20-25 percent. Development of processing and value-added products. Volume 1. Cirrhinus cirrhosus, an Indian Carp species.|Cirrhinus mrigala, an Indian Carp species. Manila, Asian Development Bank & Bangkok, Network of Aquaculture Centres in Asia-Pacific. Shrimp and carp aquaculture and the environment. Mrigal is widely cultured along with catla and rohu. Feeding and feed management of Indian major carps in Andhra Pradesh, India. Roberts (1997) considered this species as a synonym of Bangana ariza, however, this synonymy is not considered valid (Jayaram 1999, Menon 1999). The types are: 1. The Freshwater Fishes of India, Pakistan, Bangladesh, Burma and Sri Lanka. It is a bottom-feeding fish and often referred to as a scavenger. 2004. Shrimp and carp aquaculture and the environment. Jhingran, V.G. • Three day old hatchlings, measuring about 6 mm Hindustan Publishing Corporation (India), Delhi, India. Sustainable freshwater aquaculture in India. Central Institute of Fisheries Education, Mumbai, India. Gupta, S.D., Rath, S.C., Dasgupta, S.D. When synthetic formulations are used, a single dose of 0.4-0.5 ml/kg BW (females) or 0.2-0.3 ml/kg (males) is administered. Heteropneustes fossilis 8. Mrigal - Cirrhinus mrigala (Hamilton, 1822) [Cyprinidae] Taxonomic and biological features: Distinguishing characters Body elongate with depressed and obtusely rounded snout (Figures 1 and 2). Cirrhinus mrigala (Hamilton, 1822) Accepted Name Mrigal Animalia > Chordata > Actinopterygii > Cypriniformes > Cyprinidae > Cirrhinus > Cirrhinus mrigala (Hamilton, 1822) & Tripathi, S.D. 49-52. viz., the mrigal, Cirrhinus mrigala, the Nile tilapia, Oreochromis niloticus and the wallago, Wallago attu using histological and histochemical techniques. & Singh, W.J. Cirrhinus mrigala. More recently in India, freshwater carp culture has reached a stage termed as âaquaplosionâ or âblue revolutionâ due to innovative efforts of enterprising carp culturists, such as in the southern state of Andhra Pradesh, where vast areas of paddy fields were converted into fish ponds, since culture of Indian major carp was more profitable than traditional paddy cultivation. Cirrhina mrigala Bhuiyan, 1964. Culture of Indian major carp has been taken up on a commercial scale in recent years in the Haryana and Punjab states of India. In extensive systems, with a targeted production level of 2-3 tonnes/ha, the cost of production is about USD 0.30/kg, while the costs increase to USD 0.5-0.6/kg in semi-intensive culture, where the targeted production is 4-8 tonnes/ha. 32, FAO, Rome, Italy. 1995. The principle behind this polyculture system is the utilization of all available natural food niches and resources at different layers of the water body and at different trophic levels to achieve maximum production per unit area of water body. 2. Gopakumar, K., Ayyappan, S., Jena, J.K., Sahoo, S.K., Sarkar, S.K., Satapathy, B.B. Jena, J.K., Aravindakshan, P.K., Chandra, S., Muduli, H.K. Mrigal is cultured mainly as a component of carp polyculture systems in the ponds of India and Bangladesh, the major producing countries. Mrigal along with catla and rohu are extensively cultured in the freshwater bodies usually together in âpolycultureâ, âcomposite fish cultureâ or âmixed species cultureâ rather than âmonocultureâ. Talwar, P.K. mouth inferior and mouth thick, • Upper lips not continuous with lower lip. However, long distance transport of mrigal with other carps packed with crushed ice at 1:1 ratio in rectangular plastic crates (60 cm x 40 cm x 23 cm) in insulated vans is often practiced in India. Binomial nga ngaran; Cirrhinus cirrhosus (Bloch, 1795)Mga sinonimo; Cirrhinus chaudhryi Srivastava, 1968 Henicorhynchus horai (Lakshmanan, 1966) Cirrhinus horai Lakshmanan, 1966 Cirrhina macrops Steindachner, 1870 Mrigala buchanani Bleeker, 1860 Cirrhinus cuvierii Jerdon, 1849 Cirrhinus cauverii Jerdon, 1849 Cirrhina blochii Valenciennes, 1842 Dangila leschenaultii Valenciennes, 1842 MATERIALS AND METHODS Live 150 healthy fingerlings of Cirrhinus mrigala of mean weight 4.502 grams were collected from Manjri Fish Farm in Hadapsar using net and brought to the laboratory where they were cleansed with dilute KMnO 4 Journal of Applied Aquaculture, 13:251-282. S. A. Vhanalakar. Issue The species Cirrhinus mrigala (Hamilton, 1822) is considered as valid in Eschmeyer (CofF ver. Grow-out production of carps in India. Environment: milieu / climate zone / depth range / distribution range Ecology 1999. India Study Report. It is commercially cultured in India and in the Indian sub-continent and is commonly called mrigal. Thenohanellus catlae;Myxobolus bengalensis; M. catlae; M. hosadurgensis, Aeromonas hydrophila; A. sorbia; Aphanomyces invadans, Aquatic Animal Pathogen and Quarantine Information System - AAPQIS, Database on Introductions of Aquatic Species - DIAS, FAO FishStatJ – Universal software for fishery statistical time series, Network of Aquaculture Centres in Asia-Pacific - NACA, Ulcerations; exophthalmia; abdominal distension, Destroy badly infected fish; disinfect affected ponds with 0.5 ppm solution of KMnO, Raised white plaques, often with reddish peripheral zone leading to haemorrhagic spots on body, Body scales stretch out resembling pine cone; inflammation; ulceration; exophthalmia; abdominal distension, Mould grows like cotton wool on body, penetrating into the muscle; morbid muscle rot, Fungus grows out through gill blood vessels and causes necrosis of surrounding tissues; yellow-brown discolouration & disintegration of gill tissues, Addition of quick lime (50-100 kg/ha) to affected ponds; in case of limited infections, use 3-5% NaCl bath for 5-10 minutes, or 5 ppm KMnO, Skin, fin rays & operculum covered with white spores; sick fish keep rubbing against hard substratum, Dip in 1:5000 formalin solution for 1 hour for 7-10 days or in 2% NaCl for 7-10 days; affected ponds should be disinfected with quicklime at 200 kg/ha, Invasion of parasites in skin & gill region, Weakness; emaciation; raising of scales along their posterior margins; scale loss; perforation of scales; loss of chromatophores, Reduce density; add yeast to feed (1 g/kg); 2-3% NaCl bath, Gill, fin & skin affected; excessive mucus secretion, 3-5% NaCl dip treatment for 5-10 minutes; 100 ppm formalin bath; treat affected ponds with 25 ppm formalin or 4 ppm KMnO, Black nodules due to metacercarial cysts in the host body; infects eye and causes blindness, Parasites visible to naked eye attached to head & fin rays; haemorrhagic spots found in chronic cases, Drain and dry ponds showing severe Argulus infection; short duration 5 ppm KMnO, Large red or grey shallow ulcers with necrotic areas on skin; fungus extends deep into the musculature; lesions of acute dermatitis and ulcers, 200 kg/ha quicklime or 0.1 ppm CIFAX (a CIFA, India formulation). Corpus ID: 214631320. ... infecting the gill lamellae of mrigal carp, Cirrhinus mrigala, was characterized by the 18S rRNA gene sequence. Manila, Asian Development Bank & Bangkok, Network of Aquaculture Centres in Asia-Pacific. Several factors have enhanced the status of the farming of the Indian major carps: Carps are generally cultured in a closed system that involves herbivorous species, in which organic materials are used as the principal input sources, thereby making it a generally environmentally-friendly practice. Labeo gonius 3. In all the producing countries, almost all farmed mrigal is consumed in the local market. Abdomen rounded, body depth and head length nearly equal. Synopsis of Biological Data on Rohu, Cirrhinus mrigala (Hamilton, 1822). Six data classes i.e. The classification of myxosporeans is generally based on the morphology of their myxospores. Inland Fishes of India and Adjacent Countries. In: Regional Study and Workshop on Aquaculture sustainability and the Environment. 2001. Supplementary feed constitutes over 50 percent of the total input cost in carp polyculture; therefore, judicious feed management is of prime importance for enhancing profits. In India, up to 17 tonnes per hectare per year has been achieved by the progressive carp farmers employing carp polyculture practice. In India, Khan (1938) successfully induced Cirrhinus mrigala to spawn by injecting mammalian pituitary hormone. All mrigal (Cirrhinus mrigala) fry, were used in this experiment hatched from single egg mass. Somalingam, J., Maheshwari, U.K. & Langer, R.K. 1990. In: P. Keshavanath & K.V. Calcutta, India. Cirrhinus mrigala is the third most important Indian major carp, next to Catla catla (catla) and Labeo rohita (rohu). Cirrhinus macrosemion (Fowler, 1935) (synonym) Cirrhinus melanostigma Fowler & Bean, 1922 (synonym) Cirrhinus mrigala prosemion (Fowler, 1934) (synonym) Cirrhinus … The compatibility of mrigal in polyculture systems with other carps has already been established. Mysore Journal of Agricultural Sciences, 14(3):388-395. Hayat, M. 1995. A comparative study of growth rate of rohu - mrigal and mrigal - rohu hybrids and their parental species. Mrigel fish is a well-known fish of the Family Cyprinidae, belonging to the order Cypriniformes. [Draft report]. Sihag and Anuradha Bhradwaj1 Department of Zoology and Aquaculture, CCS Haryana Agriculture University, Hisar- 125 001, India Received: 20-08-2012 Accepted: 20-01-2013 ABSTRACT Diseases like epizootic ulcerative syndrome (EUS), hemorrhagic septicemia, fin and tail rot, gill Jena, J.K., Aravindakshan, P.K. Materials and Methods Madduvalas Reservoir (fig 1) is located in madduvalasa village, vangaramandal of palakonda … Isolation and Identification of Pathogenic Bacteria Associated with Diseased Fish Cirrhinus mrigala @inproceedings{N2019IsolationAI, title={Isolation and Identification of Pathogenic Bacteria Associated with Diseased Fish Cirrhinus mrigala}, author={A N. and JATT and S. A. and Tunio and S. B. and Memon and T. A. and Arain}, year={2019} } 727 pp. Bhubaneswar. Being the least preferred among the three Indian major carps, mrigal fetches comparatively low market prices, usually fetching 10-20 percent less than rohu or catla. Radhakrishnan (eds. Furthermore, the compatibility of mrigal in polyculture systems with regard to habitat preference and feeding habits is good. 86870) following Menon (1999: Ref. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. & Nayak, P.K. Cultured Aquatic Species Information Programme, Main producer countries of Cirrhinus mrigala (FAO Fishery Statistics, 2006). Different methods are used for induced breeding. ), Sustainable Indian Fisheries, pp. The fry were acclimated to laboratory condition in two cement tanks (80 l Cirrhina mrigala Shaw and Shebbeare, 1937. Cirrhinus mrigala is the third most important Indian major carp, next to Catla catla (catla) and Labeo rohita (rohu). Their body is covered with cycloid scales, and there are no scales on their head and snout blunt. 475 pp. Wallago attu 6. National Freshwater Aquaculture Development Plan.CIFA, Bhubaneswar, India, 75 pp. [Draft report]. Oxford and IBH Publishing Co. Pvt. 1995. Basavaraju, Y. The dietary supplementation of synbiotic in Cirrhinus mrigala juvenile (with initial body weight ranging from 2.87 ± 0.01 g to 3.26 ± 0.05 g) was evaluated in terms of changes in innate immunity, antioxidant activity and disease resistance against Aeromonas hydrophilla infection. ADVERTISEMENTS: The following points highlight the seventeen main types of fresh water edible fishes found in India. Generate a print friendly version containing only the sections you need. Application of the principles of Article 9 of the FAO Code of Conduct for Responsible Fisheries would be appropriate. Ayyappan, S. & Jena, J.N. CIFA. 1991. α & D. V. Muley. Forty‐four primer sequences available for four cyprinid fishes were tested to amplify microsatellite loci in Indian major carp, Cirrhinus mrigala .A total of 12 loci were successfully amplified with clear scorable patterns and five thereof were polymorphic. Channa Striatus 11. Governmental regulations and controls over the domestic marketing system are almost non-existent in these countries and demand and supply thus influence product value. Catla Catla 4. It is therefore necessary that the practicing countries should formulate guidelines and impose strict regulatory measures for the judicious use of these critical inputs. In general, carp are low-valued species fetching market prices of less than USD 1/kg at the producers' level; therefore, the use of major inputs such as seed, fertilizers and supplementary feed, besides labour costs, is kept to a minimum. Labeo Rohita 2. Their mouth is broad, transverse (the upper lip entire and not continuous with lower lip). This species and the Mrigal carp (Cirrhinus cirrhosus) are both considered distinct. Lateral line continuous to centre of caudal fin base (Figure 1). Convolutional Neural Network (CNN) is one of the most widely used architectures for image classification based on visual features. 99 pp. 1995. (Cirrhinus mrigala) •Body is elongated, depressed with rounded snout. (2005): “The Indian major carp species [Catla catla, Cirrhinus cirrhosus, and Labeo rhoita] are known However, increased emphasis on intensification for enhancing production in recent years has resulted in increased use of chemical fertilizers, feeds, therapeutics, drugs, chemicals, etc., which pose some concern. Shrimp and carp aquaculture and the environment. Synonyms: Cyprinus mrigala Hamilton, 1822. In some cases antibiotics and other pharmaceuticals have been used in treatment but their inclusion in this table does not imply an FAO recommendation. 83-133. Channa punctatus 10. Ctenopharyngodon idella (Grass carp), Cyprinus carpio (Common carp), Cirrhinus mrigala (Mori), Labeo rohita (Rohu), Hypophthalmichthys molitrix (Silver carp), and Catla (Thala), containing 915 images, popular for fish farming in the tropical areas of world including Pakistan. Myanmar Study Report. Hatchlings of mrigal normally remain in the surface or sub-surface waters, while fry and fingerling tend to move to deeper water. They have pharyngeal teeth in 3 rows 5.4.2/2.4.5 pattern; lower jaw with a small post-symphysial knob or tubercle. Export to South-East Asian and Middle-East countries. FAO Fisheries and Aquaculture Technical Paper No. & Jhingran, A.G. 1991. Veterinarski Arhiv, 65(5):143-148. When pituitary extract is used, females are injected with a stimulating dose of 2-3 mg/kg BW followed by a second dose of 5 to 8 mg/kg after a lapse of 6 hours; males are given a single dose of 2-3 mg/kg at the time of second injection of the female. However, the tendency of farmers to increase income per unit area has led to an excessive use of fertilizers, proteinaceous feeds and chemicals that may have detrimental effects on the environment. These harvesting difficulties make mrigal the least preferred species among the three Indian major carps for farmers. Mrigal is widely cultured along with catla and rohu. Bayesian length-weight: a=0.00741 (0.00462 - 0.01191), b=3.18 (3.05 - 3.31), in cm Total Length, based on LWR estimates for this species & (Sub)family-body (Ref. National Academy of Agricultural Sciences, New Delhi, India. Scales large and cycloid. Protease concentrations slightly varied but variations were prominent from species to species. It is a bottom-feeding fish and often referred to as a scavenger. Ayyappan, S. & Jena, J.K. 2003. Comparative evaluation of growth and survival of Indian major carps and exotic carps in raising fingerlings. Cirrhinus mrigala is the third most important Indian major carp, next to Catla catla (catla) and Labeo rohita (rohu). Their body is bilaterally symmetrical and streamlined. Ramakrishna, R., Shipton, T.A., Hasan, M.R.2013. Cirrhinus cirrhosus (Bloch, 1795) (mrigal carp) Cirrhinus fulungee (Sykes, 1839) (Deccan white carp) Cirrhinus jullieni Sauvage, 1878; Cirrhinus macrops Steindachner, 1870 (hora white carp) Cirrhinus microlepis Sauvage, 1878 (smallscale mud carp) Cirrhinus molitorella (Valenciennes, 1844) (mud carp) Cirrhinus mrigala (Hamilton, 1822) 17 tonnes per hectare per year has been taken up on a commercial scale in recent cirrhinus mrigala classification! Tonnes per hectare per year has been achieved by the progressive carp farmers employing carp polyculture systems in the sub-continent. Regional Study and Workshop on carp seed production, which have removed the reliance on the capture natural... Secreted by pituitary gland influences the maturation of gonads and spawning in fishes -. - rohu hybrids and their parental species mrigal is widely cultured along with the Indian sub-continent and is commonly mrigal. Ayyappan, S., Jena, J.K., Sahoo, S.K., Satapathy, B.B ) both. 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